Red-footed Booby
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The red-footed booby (''Sula sula'') is a large
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
of the
booby A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fre ...
family, Sulidae. Adults always have red feet, but the colour of the plumage varies. They are powerful and agile fliers, but they are clumsy in takeoffs and landings. They are found widely in the tropics, and breed colonially in coastal regions, especially islands. The species faces few natural or man-made threats, although its population is declining; it is considered to be a least-concern species by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN).


Taxonomy

The first formal description of the red-footed booby was by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in 1766 in the twelfth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
''. He introduced the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''Pelecanus sula''. The type locality is Barbados in the West Indies. The present genus ''Sula'' was introduced by the French scientist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
in 1760. The word ''Sula'' is Norwegian for a gannet. There are three subspecies: * ''S. s. sula'' (Linnaeus, 1766) – Caribbean and southwest Atlantic islands * ''S. s. rubripes''
Gould Gould may refer to: People * Gould (name), a surname Places United States * Gould, Arkansas, a city * Gould, Colorado, an unincorporated community * Gould, Ohio, an unincorporated community * Gould, Oklahoma, a town * Gould, West Virginia, ...
, 1838 – tropical Pacific and Indian Oceans * ''S. s. websteri''
Rothschild Rothschild () is a name derived from the German ''zum rothen Schild'' (with the old spelling "th"), meaning "with the red sign", in reference to the houses where these family members lived or had lived. At the time, houses were designated by sign ...
, 1898 – eastern central Pacific


Description

The red-footed booby is the smallest member of the booby and gannet family at about in length and with a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
of up to . The average weight of 490 adults from Christmas Island was .''CRC Handbook of Avian Body Masses, 2nd Edition'' by John B. Dunning Jr. (Editor). CRC Press (2008), . It has red legs, and its
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pla ...
and throat pouch are coloured pink and blue. This species has several morphs. In the white morph the plumage is mostly white (the head often tinged yellowish) and the
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
s are black. The black-tailed white morph is similar, but with a black tail, and can easily be confused with the
Nazca Nazca (; sometimes spelled Nasca; qu, Naska) is a city and system of valleys on the southern coast of Peru. It is also the name of the largest existing town in the Nazca Province. The name is derived from the Nazca culture, which flourished in ...
and masked boobies. The brown morph is overall brown. The white-tailed brown morph is similar, but has a white belly, rump, and tail. The white-headed and white-tailed brown morph has a mostly white body, tail and head, and brown wings and back. The morphs commonly breed together, but in most regions one or two morphs predominates; for example, at the
Galápagos Islands The Galápagos Islands ( Spanish: , , ) are an archipelago of volcanic islands. They are distributed on each side of the equator in the Pacific Ocean, surrounding the centre of the Western Hemisphere, and are part of the Republic of Ecuad ...
, most belong to the brown morph, though the white morph also occurs. The sexes are similar, and juveniles are brownish with darker wings, and pale pinkish legs, while chicks are covered in dense white down. The species has been recorded three times from
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
. In March 2016, footage of a red-footed booby being caught and killed by a
coconut crab The coconut crab (''Birgus latro'') is a species of terrestrial hermit crab, also known as the robber crab or palm thief. It is the largest terrestrial arthropod in the world, with a weight of up to . It can grow to up to in width from the tip ...
was recorded on the
Chagos Archipelago The Chagos Archipelago () or Chagos Islands (formerly the Bassas de Chagas, and later the Oil Islands) is a group of seven atolls comprising more than 60 islands in the Indian Ocean about 500 kilometres (310 mi) south of the Maldives arc ...
. After the coconut crab killed the bird, five others were observed being drawn to the site, where they competed over the meat. In September 2016, a male red-footed booby was found washed up on a beach in
East Sussex East Sussex is a ceremonial and non-metropolitan county in South East England on the English Channel coast. It is bordered by Kent to the north and east, West Sussex to the west, and Surrey to the north-west. The largest settlement in East ...
, UK, 5,000 miles from its nearest usual habitat. It was the first of its species ever recorded in the UK. The bird, later named Norman, was exhausted and malnourished. He was brought back to health before being transported by plane to an environmental center in the
Cayman Islands The Cayman Islands () is a self-governing British Overseas Territory—the largest by population in the western Caribbean Sea. The territory comprises the three islands of Grand Cayman, Cayman Brac and Little Cayman, which are located to the ...
in December 2016, where he subsequently died before ever being released into the wild. As a result of Norman's appearance, the red-footed booby was formally added to the British List by the British Ornithologists' Union on 16 August 2017. In January 2017, a red-footed booby was sighted on the
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
mainland for the first time.


Breeding

This species breeds on islands in most tropical oceans. When not breeding it spends most of the time at sea, and is therefore rarely seen away from breeding colonies. It nests in large colonies, laying one chalky blue egg in a stick nest, which is incubated by both adults for 44–46 days. The nest is usually placed in a tree or bush, but rarely it may nest on the ground. It may be three months before the young first fly, and five months before they make extensive flights. Red-footed booby pairs may remain together over several seasons. They perform elaborate greeting rituals, including harsh squawks and the male's display of his blue throat, also including short dances.


Diet

Red-footed boobies dive into the ocean at high speeds to catch prey. They mainly eat small
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
(such as flying fish) or
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fittin ...
which gather in groups near the surface.


Conservation

The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) lists the red-footed booby as a species of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
, though the population worldwide is decreasing. The warm phase ( El Niño) of the El Niño–Southern Oscillation in 1982 and 1983 negatively affected breeding on Christmas Island as higher water temperatures reduced food supply. Where usually 6000 pairs nested, 30 pairs and the around 60 pairs attempted breeding in 1982 and 1983 respectively.


References

* * Hilty. ''Birds of Venezuela''. * {{Taxonbar, from=Q217122 red-footed booby Birds of French Polynesia Birds of the Caribbean Birds of Hawaii Birds of Palau Birds of the Dominican Republic Birds of Haiti Birds of Puerto Rico Birds of Trinidad and Tobago Birds of Ascension Island Birds of the Cayman Islands Birds of the British Virgin Islands Birds of the Atlantic Ocean Birds of the Indian Ocean Birds of the Pacific Ocean Galápagos Islands coastal fauna Natural history of the Northwestern Hawaiian Islands red-footed booby red-footed booby Pantropical fauna